346. extinct because it has not been seen since 1960 [22], making S. Subscribe. [1] Una ning gihulagway ni William Chapman Hewitson ni adtong 1852. Just a moment. 700 espécies. The subfamily is considered basal within the diverse family Nymphalidae, and libytheines retain many plesiomorphic features, such as a functional female foreleg and smooth larva. No other sex. Scientific Name: Morpho peleides. Malaysian blue clipper butterfly ( - also see below and integrate. Nur-Print, Almaty. Mar 19. On the basal areas of the upperside of the wings (especially the hindwings) are large suffused patches of metallic green, blue, orange or red. Nearly all lepidopteran larvae are called caterpillars. A phylogenetic approach was used to study color pattern evolution in Vanessa butterflies. Most species also. Lymanopoda insula. The Nymphalinae are a subfamily of brush-footed butterflies ( family Nymphalidae). Nymphets are supposed to be charming,. Extant and fossil genera and species in the Libytheinae (Nymphalidae) are revised and reviewed. Hipparchia autonoe (Esper, 1783) is a protected butterfly species found in Mt. It is avoided by most insect predators. に従い、本種の学名を Erebia neriene と. the subfamily Nymphalinae, and three outgroup species from other subfamilies (Charaxinae, Limenitidinae, and Satyrinae) within the family Nymphalidae (Wahlberg et al. The mycalesine butterfly Bicyclus anynana, the "Squinting bush brown," is a model organism in the study of lepidopteran ecology, development, and evolution. Hyalyris antea achuar. Extremely Filthy Nymphets 2: With Mathilda, Suzy. We have determined mitochondrial genome of H. Contributed by Troy Bartlett on 16 February, 2004 - 12:32pm. To date, little is known about the genomic and molecular shifts associated with the evolution of the saprophagous feeding strategy. Some authorities consider this group to be at the family level (Danaidae). 200K Followers, 191 Following, 50 Posts - See Instagram photos and videos from Sabrina Oliveira (@sayumebby)Understanding ecological requirements of endangered species is a primary precondition of successful conservation practice. Class: Insecta. Sometimes, the subfamilies Limenitidinae, and Biblidinae are included here as subordinate tribe (s), while the tribe Melitaeini is occasionally regarded as a. idalia, Boloria selene, and B. Methoprene seems to terminate reproductive diapause, although transplantation. T1 - New Reports that Monarch Butterflies (Lepidoptera. [4] Male wingspans are 35–40 mm and female. Nymphalidae sp. Euphydryas cynthia. The mitogenomes of H. Diadem, Hypolimnas misippus (Linnaeus, 1764) Canary Islands, Madeira. Kupu-kupu dalam famili ini biasanya berukuran sedang hingga besar, serta mempunyai sepasang tungkai depan yang agak. 2 B), the genera Symbrenthia and Mynes were not sister. Here, we present a draft genome sequence for B. The analysis showed the taxa grouping. It is named for the mineral malachite, which is similar in color to the bright green on the butterfly's wings. The wings, bodies, and legs, like those of moths, are covered with dustlike scales that come off when the animal is handled. The Junonia lemonias mitogenome maintains synteny with other Nymphalid butterflies. The mitogenomes of H. The checklist is made based on the recent updated taxonomic data and original descriptions. The undersides range from bright red to pale brown, interlaced by black and white. This study compiles previously published morphological, colour and behavioural characters and includes new DNA sequence data for eight markers (one mitochondrial and seven nuclear) to re-evaluate phylogenetic relationships and estimate times of divergence for Morpho butterflies using parsimony and Bayesian methods. This is a list of the butterflies of China belonging to the family Nymphalidae and an index to the species articles. Related Hot 3d porn videos in HD Hot 3d porn 3d hot gay porn 3d mama New 3d porn videos 3D animation hot porn vid 3d sprem hot girl porn Animation porn hot Hot sex. scott small. Satyrinae. The color patterns on the wings of butterflies and moths are among the most complex manifestations of pattern formation in nature. [1] [2] Walay nalista nga matang nga sama niini. Relationship between growth and molting periods of the same instar (from L1 to L5 plus two extra L6) for 13 Hypothyris caterpillars. You can follow me in IG: I love Mario Games, especially Mar. A powerful flier, this species is. Common wood-nymph (. At 106 tallgrass prairies in the midwestern USA, 18,055 individuals of six fritillary species (Lepidoptera: Nymphalidae) were counted in 552. They were formerly considered a distinct family, Satyridae. @Nymphelita_. They are found in the rainforests and secondary forests of Mexico, Central and South America. Nymphalinae. p. [1] Una ning gihulagway ni William Chapman Hewitson ni adtong 1874. Blue Morpho Butterfly. OnlyFans. Hübner, 1819. Hewitson, 1852. Ninfalídeos[ 1] ( Nymphalidae) é uma família de insectos da ordem Lepidoptera que inclui cerca de 6 000 espécies, entre as quais a maioria das espécies mais conhecidas e coloridas de borboletas. We offer Adel 🌷 OnlyFans leaked content, you can find a list of available content of nymphelita below. Ithomia patilla. Papilio laodamia Cramer, 1777. It includes the following species: [1] Doxocopa agathina (Cramer, [1777]) - Agathina emperor. C. This family is easily recognized by reduced front legs that lack claws. Chestnut Tiger. Belonging to the superfamily Papilionoidea, they. The complete life cycle takes 74 days at ambient. The chemical ecology of plant-insect interactions has been driving our understanding of ecosystem evolution into a more comprehensive context. nymphelita OnlyFans profile was leaked recently by anonymous. Many species, like the monarch butterfly, are very colorful. Many species, like the monarch butterfly, are very colorful. Mitotic and meiotic chromosomes from 2 taxa of the genus Melinaea, M. 7K Views. The owl butterflies are species of the genus Caligo and are known for their huge eyespots, which resemble owls ' eyes. m. 6 in) for females. The complexities of these patterns arise from the diversification of a conserved set of homologous elements known as the Nymphalid Ground Plan that can change color, shift position, expand, or disappear altogether. OnlyFans. Snout butterflies (Nymphalidae: Libytheinae) are morphologically one of the most unusual groups of Lepidoptera. In males the dorsal sides of the wings are bright orange red, framed by a black outline with white spots. The family Nymphalidae is the most speciose family of butterflies with about 6000 described species so far. The members of the Papilionoidea may be distinguished by the following combination of characters: The body is smaller and less moth-like. Molly&Linni. #Petite #braces #brazil #redhead #brunette #skinny #small boobs #small tits #cute #teen #ddlg #nymphetty #slender #adoreable. assimilis, which is typical circular duplex molecules and 15,262 bp in. Nymphalinae. Distribution. satevis cydon and M. Nymphalinae Tortoiseshells, Admirals,. 1254. 2 • INSECTA MUNDI 0353, February 2014 COCK Family Hedylidae Guenée, 1858 (1 genus, 36 species) Family Hesperiidae Latreille, 1809 (570 genera, 4,113 species) Family Pieridae Swainson, 1820 (91 genera, 1,164 species)Most butterflies feed on nectar, while some saprophagous butterflies forage on various non-nectar foods. Six species were sequenced, aligned, and used to construct phylogenetic trees. 1. Despite the fact that Bicyclus. [1] Its wingspan is approximately 12 cm (4. Nokkosperhoset, Amiraalit,. Kaliwatan sa alibangbang ang Lymanopoda insula. sita, we investigated. 99 331K. Comparison of moth and butterfly dorsal anatomy. PDF | Vanessa atalanta (Linnaeus, 1758) is reported from Tajikistan for the first time. The well known caterpillars are banded with yellow, black, and cream. nov. Order: Lepidoptera. Abstract. @Nymphelita_. 000 spesies kupu-kupu yang tersebar di seluruh dunia. Adult Essex skipper (Thymelicus lineola) Pupa of a sphingid moth Caterpillar of the subfamily Arctiinae Eggs of the buff-tip (Phalera bucephala), a notodontid moth. 17) which are capable of piercing skin. Parsimony analyses of forty-nine species in twenty-nine genera indicate that He. 2018, 9, 179 3 of 16. Species: C. Their front two legs look more like brushes than feet. The. The relationships of major lineages in Nymphalidae have been unclear for a long period, but recent work by the NSG, Andrew Brower and André Freitas are. Named for its conspicuous target-shaped eyespots, the common buckeye, Junonia coenia Hübner, is one of the most distinctive and readily-identifiable North American butterflies. 700 spesies. 50 USD. This page was last edited on 6 April 2022, at 15:30. 11. Some authorities consider this group to be at the family level (Danaidae). Ang Lymanopoda insula sakop sa kahenera nga Lymanopoda, ug kabanay nga Nymphalidae. 1093/gigascience/gix035. Butterflies in this family are called brush-foots because of their tiny forelegs. Binomial name. The mitochondrial genomes (mitogenomes) of 11 new nymphalid species were reported and a comparative mitogenomic analysis was conducted together with other 22 available nymphalid mitogenomes. zerene). This pattern is available for $6. 8% T, 15. Tribe Nymphalini. 6%). To this contribution, the complete mitogenome of Danaus genutia (. 340. モルフォチョウ （モルフォ蝶、 学名 ： Morpho ）は、 北アメリカ 南部から 南アメリカ にかけて80種ほどが生息する大型の チョウ の仲間。. Limenitis arthemis, the red-spotted purple or white admiral, is a North American butterfly species in the cosmopolitan genus Limenitis. 2008. The mourning cloak, Nymphalis antiopa (Linnaeus), is a large distinctive butterfly and is one of our most widely distributed butterfly species. Distributed worldwide, butterflies of this family are especially rich in the tropics. The phylogenetic analyses of the subfamily Limenitidinae are performed based on 1 471 bp of mtDNA cytochrome oxidase subunit I (COI) gene sequence data which were obtained from 21 individuals spanning 9 genera, along with those of 17 species obtained from GenBank, using Apatura iris, Aglais urticae, and Polyura dolon as outgroup. Nymphalidae genus: AbrotaThe case of Nymphalidae on display from the . It has a near-global distribution and includes conspicuous species such as the red admirals (e. Malaysian blue clipper butterfly ( - also see below and integrate. The viceroy was named the state butterfly of Kentucky in 1990. Many fritillary. The Libytheinae includes two genera: Libythea Fabricius and Libytheana Michener. The Nymphalidae family of cosmopolitan butterflies (Lepidoptera) comprises approximately 7200 species found on all continents and in all habitats. [2] It is primarily found in eastern Africa from southern Sudan to Eswatini. The start codons of the protein-coding genes (PCGs) in the. See examples of NYMPHALID used in a sentence. 5 in), leathery-winged butterfly common in grassland and scrub habitats. Morpho é um gênero de borboletas predominantemente azuis, classificadas na família Nymphalidae, que vivem nas florestas tropicais da América central e da América do Sul: Brasil, Costa Rica, Guiana e Venezuela mas também no México. Houghton Mifflin Company. 😍💓 ️💓 ️😘. Most species in this family are medium sized, sun-loving butterflies and are recognised by their rapid, direct flight and the tendency to settle on the ground with their brightly coloured wings outstretched. This group contains nearly half of the known diversity of brush-footed butterflies. The head of a small white butterfly (Pieris rapae). In contrast to the. Quick Facts. also available for free: info in notes. We are doing our best to renew the leaks of nymphelita. 一般に成虫は中型で寿命が長いものが多い。. беларуская: Німфаліды čeština: Babočkovití dansk: Takvingefamilien Deutsch: Edelfalter English: Brush-footed butterfly español: Ninfálidos. Euploea mulciber. Three new synonyms are proposed: Chalinga Moore, 1898 = Seokia Sibatani, 1943. Basic Insect Anatomy. Backup (@nymphelita_) Twitter. Satyrinae. Houghton Mifflin Company. Every day, thousands of people use EroMe to enjoy free photos and videos. It occurs in large populations in montane and lowland forests, typically in areas of secondary plant growth (Young 1973, Young 1975, Urich and Emmel 1991). gallardi Neukirchen, 1996 (Lepidoptera, Nymphalidae) - Asterope markii Hewitson, 1857, two new subspecies in Peru and description of the male of A. Golden Banded-Skipper ( Autochton cellus) Grizzled Skipper ( Pyrgus centaureae) Hayhurst’s Scallopwing ( Staphylus hayhurstii) Hoary Edge ( Achalarus lyciades) Horace’s Duskywing ( Erynnis horatius) Juvenal’s Duskywing ( Erynnis juvenalis) Long-tailed Skipper. 学名. This is a list of the butterflies of China belonging to the family Nymphalidae and an index to the species articles. The superfamily Papilionoidea (from the genus Papilio, meaning "butterfly") contains all the butterflies except for the moth-like Hedyloidea . . Replying to. The Nymphalidae Systematics Group is now part of the Systematic Biology Group at Lund University. Binomial name. This pair of legs is . A Catalogue of. It is either considered a sister group of the Satyrinae, or disassembled and included therein. dan rujuk teks rencana. Family NYMPHALIDAE is the largest butterfly family, with sizes that vary from large to small, most are in medium size. pegala. No other sex tube is more popular and features more Onlyfans Nymphelita scenes than Pornhub! The Nymphalidae are the largest family of butterflies, with more than 6,000 species distributed throughout most of the world. ( ˈnɪmfəlɪd) n. The antennae are straight and clubbed or. Insects. The circular mitogenome of H. Perhaps the most distinctive physical features of the butterfly are its club-tipped antennae and its habit of holding the. Hotleak is the best free porn site of Millions Exclusive Leak contents such as Images, Gallery and Videos. Whereas these American limenitidines have received the most attention, phylogenetic relationships of their Asian. Parthenos sylvia, the clipper, [2] [3] is a species of nymphalid butterfly found in south and southeast Asia, mostly in forested areas. Bicyclus anynana ( squinting bush brown) is a small brown butterfly in the family Nymphalidae, the most globally diverse family of butterflies. Diversitas. n. Insects. , 2020). We investigated the effects of juvenile hormone analogue (methoprene) and 20-hydroxyecdysone on female and male reproduction in a nymphalid butterfly, Polygonia c-aureum. The external. They have a velvety, black and speckled body filled with hairy of spiky projections, alongside small, white dots. My only account (@sayumebby) TikTok. The site is inclusive of artists and content creators from all genres and allows them to monetize their content while developing authentic relationships with their fanbase. The Satyrinae, the satyrines or satyrids, commonly known as the browns, are a subfamily of the Nymphalidae (brush-footed butterflies ). The front legs, due to their length, are not used for walking. Adults are delicate and fly close to the ground in a slow, bouncy flight pattern that is characteristic of butterflies from the subtribe Euptychiina. The Hestina assimilis belongs to Nymphalidae in Lepidoptera. Both sexes have the same appearance, and the wingspan is 3 to 4 inches. The Menelaus blue morpho ( Morpho menelaus) is one of thirty species of butterfly in the subfamily Morphinae. The origin and dispersal history of the large butterfly subfamily Nymphalinae are not fully understood, due to internal phylogenetic and time calibration issues. Hola 😘🌹 Eres bonita y admirable. l-album, the Compton tortoiseshell, or false comma, is a species of butterfly in the family Nymphalidae. Woon-Young Chun from Hainan Island, China (Lepidoptera, Papilionoidea), Zootaxa: Vol. Use the By Field search to find specimen data that match values in specific database fields. The site is inclusive of artists and content creators from all genres and allows them to monetize their content while developing authentic relationships with their fanbase. Nymphalidae: Adelpha californica. Just a moment. Occasional hosts in Trinidad are also Erythrina glauca and E. Instead,. Lepidopteran, insect order containing numerous species of butterflies, moths, and skippers. 2 A), S. adiaste the only Speyeria species to have a described subspecies go extinct (but see [34] for discussion of S. florissanti, and L. 1: 14 Feb. The mourning cloak, Nymphalis antiopa (Linnaeus), is a large distinctive butterfly and is one of our most widely distributed butterfly species. g. A generic-level phylogeny for the butterfly family Nymphalidae was produced by cladistic analysis of 234 characters from all life stages. Estimates of inbreeding consequences were obtained by comparing 12 life-history and morphological traits among nine inbred families (F = 0. 2023. Binomial name. Nymphalidae in Charles Oberthür's Études d'entomologie: Lépidoptères de Chine (1876). It has a weak fluttery flight. We study the phylogenetic relationships of taxa in the butterfly family Nymphalidae as well as other families of Lepidoptera,. PDF | Vane-Wright, R. These are usually medium-sized to large butterflies. 16 spp. They live on all continents except Antarctica [1] They are commonly known as nymphalids, brushfoots, or brush-footed butterflies . A Field Guide to Eastern Butterflies (Peterson Field Guides) Paul A. Cerca de 5. Indeed, nymphalids are in many places the most visible members of the local. My only account (@sayumebby) TikTok. Hyalyris antea cana (Haensch, 1905) TL: Colombia M T. List of butterflies of India (Nymphalidae) Clipper ( Parthenos sylvia ), a nymphalid butterfly found in India. Abstract and Figures. The Nymphalidae is a family of several thousand species found in all zoogeographical regions of the world. Comparison of moth and butterfly dorsal anatomy. We here propose a new, monotypic genus, Amiga Nakahara, Willmott & Espeland, gen. Nymphetty | Nymphelita | Sayume_Mii pictures and videos on EroMe. Parsimony analyses used both equal weights and implied weights, and a series of analyses were performed. sita, we investigated both molecular data and morphological features in 429. Enter a value or choose one from the dropdown lists. Request PDF | A new species related to Ypthima praenubila Leech, 1891 in Vietnam and China (Lepidoptera, Nymphalidae, Satyrinae) | We describe a new species of satyrine butterfly, Ypthima thao. Main page (@sayumebby)The Carolina satyr, Hermeuptychia sosybius (Fabricius, 1793) has a range that spans 13 states across the southeastern US and is often abundant where it occurs. The Nymphalidae family of butterflies are a family of medium to large sized, brightly colored butterflies. It was long thought to be a Batesian mimic of the monarch butterfly, but since the viceroy is also distasteful to predators, it is now considered a Müllerian mimic instead. A free version is also available. Recent studies on the ecology and genetics of speciation in Ithomiini have. It spends much of its time in second-growth woodlands. They dominate by number the mimetic butterfly communities, which include species such as the iconic neotropical Heliconius genus. THE NCBI Taxonomy database allows browsing of the taxonomy tree, which contains a classification of organisms. Kaliwatan sa alibangbang ang Lymanopoda cillutinarca. , the residuals of the linear regression of thoracic mass on body mass, [10]) and flight muscle ratio (FMR = thoracic mass. Although a number of lineages within the family are well circumscribed and have been recognized since the 1800s, the taxonomic rank which such lineages should be accorded and how they should be subdivided. January 27, 2022 by Active Wild Admin. Home Games & Quizzes History & Society Science & Tech Biographies Animals & Nature Geography & Travel Arts & Culture Money Videos. We assembled the dataset with sampling the largest number of species and genera of this subfamily so far (comprising one mitochondrial and two nuclear gene sequences) to conduct. Nymphalidae. Hyalyris antea (Hewitson, 1869) Hyalyris antea anteaTL: Ecuador. The life history of Morpho peleides insularis (Fruhstorfer) (Lepidoptera: Nymphalidae: Morphinae) is described from material reared on Paragonia pyramidata (L. Opler, Vichai Malikul, Roger Tory Peterson. Malay Lacewing ( Cethosia hypsea hypsina. The Nymphalidae family of cosmopolitan butterflies (Lepidoptera) comprises approximately 7200 species found on all continents and in all habitats. In addition to three pairs of legs on the thorax, they have two to eight pairs of fleshy abdominal prolegs that are structurally different from the. 3–3. Limenitidinae. Most species have a reduced pair of forelegs and many hold their colourful wings flat when resting. Mitogenomic phylogenies have revealed well-supported relationships for many eukaryote groups. Twenty‐four color pattern elements from the Nymphalid ground plan were identified on the dorsal and ventral surfaces of the fore‐ and hind wings. . Hypolimnas misippus, a sexually dimorphic, nymphalid butterfly with a very variable female, was sampled for 41 consecutive months, along with its supposed model, Danaus chrysippus (Danaidae), at Dar es Salaam, Tanzania. Ang Lymanopoda cillutinarca sakop sa kahenera nga Lymanopoda, ug kabanay nga Nymphalidae. The sole representative of its genus Dryas, it is native from Brazil to southern Texas and Florida, and in summer can sometimes be found as far north as eastern Nebraska. plexippus found in North America. A. The name Limenitis is Neo-Latin "of harbours", from Ancient Greek Λιμενιτις (from λιμήν, a harbour, haven). 25) and 16 outbred families (F = 0) of the Neotropical butterfly Heliconius erato phyllis. In northern areas where it overwinters, adults may be seen basking in the sun during almost every month of winter on warm days. Watch and download Free OnlyFans Exclusive Leaked videos of Nymphelita 🌷 [ nymphelita ] in high quality. v. Lepidoptera (moths and butterflies) is the second largest order in the class Insecta. . The common names of many species and genera reference military ranks or – namely the Adoliadini – titles of nobility (e. Nymphalidea is named for one of the largest families of butterflies, with varieties found in every corner of the world. @Nymphelita_. If you are interested in more. Subfamily: Nymphalinae. The analysis showed the taxa grouping into six main lineages. Mais de 600 gêneros. [1] [2] Walay nalista nga matang nga sama niini. It belongs to the Nymphalidae or brush-footed butterfly family. Subfamily Nymphalinae. In this paper, we describe and illustrate the adult. 2018, 9, 179 3 of 16. The higher-level classification of this family is still controversial. Status: Widespread common butterfly, best known as migrant. Species recognized are: Libythea celtis (Laicharting), L. Jonathan P. The circumscription of Nymphalinae has enjoyed a dynamic history as various authors have. from northeastern Brazil, based on morphological and molecular data. PDF | Three new species and seven new subspecies of Philaethria from Colombia, Venezuela, and Panama are described as follows: P. A Student's t-test. vagabunda. Sekitar 5. buy it now or visit pattern website. The fauna of montane satyrine butterflies of the subtribe Pronophilina (Lepidoptera: Nymphalidae) is surveyed in the Venezuelan Cordillera de la Costa, a long but relatively low range of mountains consisting of two parallel units, the northern and higher Serranía del Litoral and the southern, lower Serranía del Interior. 99 323,8K. Nymphalidae: Adelpha californica. 本種の分類にかんしては議論があるが、本項では 日本産蝶類和名学名便覧 2010-2013. Hypodryas cynthia Higgins, 1981. Animals that start with X include: mammals such as Xanthippe’s shrew and the xolo (a breed of dog); the reptile Xantus’ leaf-toed gecko (a species of lizard); fish such as the X. Lebih dari 600 genus. Southern regions of the US, especially Florida and Texas, and also ranges southward through Mexico; they have two migrations during the spring and the fall, respectively. Hamadryas laodamia, the starry night cracker or starry cracker, is a species of cracker butterfly in the family Nymphalidae. 7K Views. The base composition of the COI sequences was 37. The Libytheinae includes two genera: Libythea Fabricius and Libytheana Michener. The site is inclusive of artists. It is now clear that there are 12 subfamilies that are well. ベニヒカゲ. ( Cramer, 1777) Synonyms. Introduction. sita using a species distribution model (Maxent),. Persius Duskywing (. To clarify the level of differentiation within P. Tarucus balkanicus (Freyer, 1843) Zizeeria karsandra (Moore, 1865) Brephidium exilis (Boisduval, 1852) Chilades parrhasius (Fabricius, 1793)Here are 12 blue butterfly species you should look into if you’re curious about the different types of blue butterflies: 1. Most are medium or large in size, but the family is highly variable given that it also includes the Satyrinae, a subfamily that has been designated as a family in its own right in earlier classifications. be careful with fake profiles, I don't have private pages to talk. quick link to butterfly pages. Lymanopoda cillutinarca. The majority of species are found in both Old and New World tropics. 2014 SHAO-JI HU, FABIEN L. Background Major conflict between mitochondrial and nuclear genes in estimating species relationships is an increasingly common finding in animals. 17 539. @Sayume_Mi sooooooooo adorable princess/queen 😘😘😘. Backup (@nymphelita_) Twitter. With around 6,000 species, this is the largest family of butterflies. White peacock butterfly ( Anartia jatrophae. micropteryx (Fabaceae). quick link to butterfly pages. PDF | On Dec 19, 2018, Vitthalrao Bhimasha Khyade and others published Explanation of Nymphalidae Butterflies | Find, read and cite all the research you need on ResearchGate Nymphalidae: [plural noun] a widely distributed family of butterflies mostly of medium or large size and distinguished by having the forelegs much reduced in size in both sexes so that they are useless in walking and are carried folded on the breast, usually absent tarsal claws, larvae that are usually spiny or provided with fleshy processes,. , P,. DOI: 10. Ancestral range and historical biogeographic events were reconstructed using the Bayesian binary MCMC method (Ronquist and Huelsenbeck, 2003) and BioGeoBEARS (Matzke, 2013a, Matzke, 2013b) implemented in RASP (Yu et al.